Evaluating revised biomass equations: are some forest types more equivalent than others?
- Coeli M. Hoover^{1}Email author and
- James E. Smith^{1}
https://doi.org/10.1186/s13021-015-0042-5
© Hoover and Smith. 2016
Received: 26 October 2015
Accepted: 22 December 2015
Published: 12 January 2016
Abstract
Background
In 2014, Chojnacky et al. published a revised set of biomass equations for trees of temperate US forests, expanding on an existing equation set (published in 2003 by Jenkins et al.), both of which were developed from published equations using a meta-analytical approach. Given the similarities in the approach to developing the equations, an examination of similarities or differences in carbon stock estimates generated with both sets of equations benefits investigators using the Jenkins et al. (For Sci 49:12–34, 2003) equations or the software tools into which they are incorporated. We provide a roadmap for applying the newer set to the tree species of the US, present results of equivalence testing for carbon stock estimates, and provide some general guidance on circumstances when equation choice is likely to have an effect on the carbon stock estimate.
Results
Total carbon stocks in live trees, as predicted by the two sets, differed by less than one percent at a national level. Greater differences, sometimes exceeding 10–15 %, were found for individual regions or forest type groups. Differences varied in magnitude and direction; one equation set did not consistently produce a higher or lower estimate than the other.
Conclusions
Biomass estimates for a few forest type groups are clearly not equivalent between the two equation sets—southern pines, northern spruce-fir, and lower productivity arid western forests—while estimates for the majority of forest type groups are generally equivalent at the scales presented. Overall, the possibility of very different results between the Chojnacky and Jenkins sets decreases with aggregate summaries of those ‘equivalent’ type groups.
Keywords
Biomass estimation Allometry Forest carbon stocks Tests of equivalence Individual-tree estimates by species groupBackground
Nationally consistent biomass equations can be important to forest carbon research and reporting activities. In general, the consistency is based on an assumption that allometric relationships within forest species do not vary by region. Essentially, nearly identical trees even in distant locations should have nearly identical carbon mass. In 2003, Jenkins et al. published a set of 10 equations for estimating live tree biomass, developed from existing equations using a meta-analytical approach, which were intended to be applicable over temperate forests of the United States [1]. These equations were developed to support US forest carbon inventory and reporting, and had several key elements: (1) a national scale, so that regional variations in biomass estimates due to the use of local biomass equations was eliminated, (2) the exclusion of height as a predictor variable, and (3) in addition to equations to estimate aboveground biomass, a set of component equations allowing the separate estimation of biomass in coarse roots, stem bark, stem wood, and foliage. Since their introduction, these equations have been incorporated into the Fire and Fuels Extension of the Forest Vegetation Simulator as a calculation option [2], utilized in NED-2 [3], and have provided the basis for calculating the forest carbon contribution to the US annual greenhouse gas inventories for submission years 2004–2011 (e.g., see [4]). Researchers in Canada [5, 6] and the US (e.g. [7–9]) have also employed the equations while other investigators have adopted the component ratios to estimate biomass in coarse roots or other components (e.g. [10, 11]).
In 2014, Chojnacky et al. [12] introduced a revised set of generalized biomass equations for estimating aboveground biomass. These equations were developed using the same underlying data compilations and general approaches to developing the individual tree biomass estimates as for Jenkins et al. [1], but with greater differentiation among species groups, resulting in a set of 35 generalized equations: 13 for conifers, 18 for hardwoods, and 4 for woodland species. Important distinctions are: the database used to generate the revised equations was updated to include an additional 838 equations that appeared in the literature since the publication of the 2003 work or were not included at that time, taxonomic groupings were employed to account for differences in allometry, and taxa were further subdivided in cases where wood density varied considerably within a taxon. The only component equation revised by Chojnacky et al. [12] was for roots; equations were fitted for fine and coarse roots, in contrast to Jenkins et al. [1] where fine roots were not considered separately.
Based on the similarity of the equation development approach, it is likely that applications using the Jenkins et al. [1] set would have essentially the same basis for employing the revised equations. Since the primary objective of Chojnacky et al. [12] was to present the updated equations and describe the nature of the changes, only a brief discussion of the behavior of the updated equations vs. the Jenkins et al. [1] equation set was included. The authors noted that at a national level results were similar, while differences occurred in some species groups, for example, western pines, spruce/fir types, and woodland species. Given the limited information provided in Chojnacky et al. [12] we felt that a more thorough investigation of the differences in carbon stock estimates as generated with both sets of equations was needed.
One potentially practical result from a comparison of the two approaches is to identify where one set effectively substitutes for the other, which then suggests that revising or updating estimates would change little from previous analyses. For this reason we applied equivalence tests to determine the effective difference of the Chojnacky-based estimates relative to the Jenkins values. Note that hereafter we label the respective equations and species groups as Chojnacky and Jenkins (i.e., in reference to their products not the publications, per se).
In this paper, we: (1) provide a roadmap for applying the Chojnacky equations to the tree species of the US Forest Service’s forest inventory [13], (2) present results of equivalence testing for carbon stock estimates computed using both sets of equations, and (3) provide general guidance on the circumstances when the choice of equation is likely to have an important effect on the carbon stock estimate. Note that we do not attempt any evaluation of relative accuracy or the relative merit of one approach relative to the other.
Results and discussion
Forest type group | All US^{a} | North | South | Rocky Mountain | Pacific Coast | |||||
---|---|---|---|---|---|---|---|---|---|---|
Jenkins | Chojnacky | Jenkins | Chojnacky | Jenkins | Chojnacky | Jenkins | Chojnacky | Jenkins | Chojnacky | |
White/red/jack pine | 68.7** | 67.2** | 67.7** | 66.2** | 92.4** | 93.5** | ||||
Spruce/fir | 45.8 | 40.1 | 47.5 | 41.6 | 20.5* | 18.9* | ||||
Longleaf/slash pine | 35.4 | 40.6 | 35.4 | 40.6 | ||||||
Loblolly/shortleaf pine | 47.0 | 54 | 59.0 | 67.1 | 47.2 | 54.1 | ||||
Pinyon/juniper | 18.4 | 22.5 | ^{◊}15.5 | ^{◊}17.2 | 11.5 | 13.3 | 19.6 | 24.1 | 21.4 | 23.4 |
Douglas-fir | 114.5* | 108.0* | 71.4* | 66.5* | 148.6* | 140.9* | ||||
Ponderosa pine | 50.0** | 50.7** | 37.3** | 37.9** | 46.3** | 47.1** | 53.5** | 54.2** | ||
Western white pine | 66.2** | 67.6** | ^{◊}74.6 | ^{◊}76.7 | ||||||
Fir/spruce/mtn hemlock | 92.2* | 87.1* | 71.8 | 64.4 | 119.4** | 117.4** | ||||
Lodgepole pine | 48.6** | 48.2** | 48.2** | 47.2** | 49.5** | 49.7** | ||||
Hemlock/sitka spruce | 155.1** | 151.0** | 108.8* | 101.4* | 159.7** | 155.9** | ||||
Western larch | 62.6** | 65.2** | 55.4** | 57.5** | 69.6 | 72.6 | ||||
Redwood | 236.2** | 235.3** | 236.2** | 235.3** | ||||||
Other western softwoods | 27.0 | 35.3 | 43.2 * | 45.8 * | 19.5 | 30.4 | ||||
California mixed conifer | 134.7** | 132.8** | 134.7** | 132.8** | ||||||
Oak/pine | 54.1** | 56.6** | 64.4** | 65.5** | 50.9 * | 53.9 * | ||||
Oak/hickory | 72.7** | 72.8** | 78.7** | 78.8** | 65.2** | 65.3** | ||||
Oak/gum/cypress | 78.1** | 79.7** | 86.9** | 85.2** | 78.5** | 80.3** | ||||
Elm/ash/cottonwood | 56.6** | 56.6** | 60.6** | 59.8** | 50.4** | 52.2** | 48.8** | 48.2** | 82.3 | 71.8 |
Maple/beech/birch | 80.7** | 80.3** | 80.1** | 79.7** | 82.1** | 83.3** | ||||
Aspen/birch | 45.3** | 43.2** | 43.9** | 41.8** | 52.8** | 50.4** | 38.0** | 36.5** | ||
Alder/maple | 98.5** | 100.1** | 99.4** | 101.0** | ||||||
Western oak | 64.7* | 61.1* | 64.7** | 61.1** | ||||||
Tanoak/laurel | 131.2** | 134.6** | 131.2** | 134.6** | ||||||
Other hardwoods | 49.6** | 51.2** | 43.0* | 45.8* | 43.2* | 45.9* | 67.5** | 66.3** | ||
Woodland hardwoods | 8.6 | 11.1 | 5.0 | 7.0 | 12.7 | 15.7 | 22.1 | 29.5 |
Pinyon/juniper was not equivalent in any region in which it was tested. While fir/spruce/mountain hemlock was not equivalent in the Rocky Mountains, the stock estimates were equivalent to 5 % in the Pacific Coast region, likely a function of the species and size classes that dominate the groups in each of these regions. The elm/ash/cottonwood category is represented in each region, and was equivalent to 5 % in all areas except the Pacific Coast. The woodland class has been less well studied than the others, and so less data and fewer equations are available to construct generalized equations like those in Jenkins et al. [1] and Chojnacky et al. [12]. Consequently, the woodland equations are not equivalent at the national level or in any region.
Conclusions
The revised approach to developing these biomass equations has the effect of providing better regional differentiation/representation at the plot/stand level summaries by allowing for separation within the taxonomic classes according to wood properties or growth habit. The emergence of Southern pines as distinctly different under the Chojnacky groups is one example. It is challenging to provide specific criteria for choosing one set of equations over the other, since validating any biomass equation requires the destructive sampling of multiple stems across a range of diameters. The Chojnacky groups appear to provide greater resolution across forest types and regions. From this, investigators working in southern pine, northern spruce-fir, pinyon-juniper, and woodland types may be advised to use the updated equations [12], which provide more taxonomic resolution. It should also be noted that estimates of change over time are somewhat less sensitive to equation choice than stock estimates, so if change is the primary variable of interest, the user can select either equation set, based on personal preference.
Individual large diameter trees can be very different—Chojnacky relative to Jenkins—given the general trends of the tree-level estimates (Figs. 4 and 5 in this manuscript as well as Figs. 2, 3, and 4 in Chojnacky et al. [12]). This effect of one or a very few larger trees can result in very different estimates even in an “equivalent” forest type group, and this potential for larger differences is reflected in plot-level data. For example, in some eastern hardwood type groups, which were consistently identified as equivalent, up to one-third of the plots were individually more than 5 % different. The oak/gum/cypress type group in the South had 8 % of the plots with greater carbon density by over 5 % with the Jenkins estimates, while 27 % of plots had over 5 % greater carbon. The remaining 65 % of the individual plots are within the 5 % bounds (data not shown here). This is consistent with our observation about similarities between the two sets and scale (Fig. 2)—the sometimes obvious and large differences for some forest type groups (all scales) become obscured when summed to total live tree carbon for the US. Singling out the correct or most accurate equations is beyond the scope here; however, caution is always warranted when applying equations to trees that are considerably outside the range of diameters used to construct the equations [16].
Our results point to a few forest type groups that are clearly not equivalent—southern pines, northern spruce-fir, and lower productivity arid western forests—while the majority of forest type groups are generally equivalent at the scales presented. Overall, the possibility of very different results between the Chojnacky and Jenkins sets decreases with aggregate summaries of those ‘equivalent’ type groups.
Methods
Tree data source
In order to implement the revised biomass equations and identify applications where they are effectively interchangeable, or equivalent, we used the Forest Inventory and Analysis Data Base (FIADB) compiled by the Forest Inventory and Analysis (FIA) Program of the US Forest Service [13]. The data are based on continuous systematic annualized sampling of US forest lands, which are then compiled and made available by the FIA program of the US Forest Service [18]; the specific data in use here were downloaded from http://apps.fs.fed.us/fiadb-downloads/datamart.html on 02 June 2015. Surveys are organized and conducted on a large system of permanent plots over all land within individual states so that a portion of the survey data is collected each year on a continuous cycle, with remeasurement at 5 or 10 years depending on the state. The portion of the data used here include the conterminous United States (i.e., 48 states), and the portion of southern coastal Alaska that has the established permanent annual survey plots (the gray areas in Fig. 1).
Our focus here is on the tree data of the FIADB, and for this analysis we present the Chojnacky and Jenkins estimates in terms of carbon mass (i.e., kg carbon per tree or tonnes per hectare per plot). We use the entire tree data table to assure that all applicable species (the gray areas in Fig. 1) are represented. All other summaries are based on the most recent (most up-to-date) set of tree and plot data available per state, with the Chojnacky and Jenkins estimates expressed as tonnes of carbon per hectare in live trees on forest inventory plots. These plot-level values are expanded to population totals, that is, total carbon stock per state, as provided within the FIADB as the basis for the result presented in Fig. 2. A subset of the current forest plot level summaries where the entire plot is identified as forested (i.e., single condition forest plots) is the basis for the results provided in Table 1 and Fig. 3.
Application of Chojnacky et al. [12] to the FIADB
Guide to applying Chojnacky species groups (as shown in Table 5, Chojnacky et al. [12]) to US species
Scientific name | Common name | Jenkins group | Chojnacky et al. parameters when diameter is measured at | |
---|---|---|---|---|
Breast height | Root collar | |||
Abies spp. | Fir spp. | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. amabilis | Pacific silver fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. balsamea | Balsam fir | T Fir/Hem | Abies; LoSG | Pinac; WL |
A. bracteata | Bristlecone fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. concolor | White fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. fraseri | Fraser fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. grandis | Grand fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. lasiocarpa var. arizonica | Corkbark fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. lasiocarpa | Subalpine fir | T Fir/Hem | Abies; LoSG | Pinac; WL |
A. magnifica | California red fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. shastensis | Shasta red fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
A. procera | Noble fir | T Fir/Hem | Abies; HiSG | Pinac; WL |
Chamaecyparis spp. | White-cedar spp. | Cedar/Larch | Cupr; MedSG | Cupre; WL |
C. lawsoniana | Port Orford cedar | Cedar/Larch | Cupr; MedSG | Cupre; WL |
C. nootkatensi | Alaska yellow cedar | Cedar/Larch | Cupr; HiSG | Cupre; WL |
C. thyoides | Atlantic white cedar | Cedar/Larch | Cupr; MedSG | Cupre; WL |
Cupressus spp. | Cypress | Woodland | Cupr; HiSG | Cupre; WL |
C. arizonica | Arizona cypress | Woodland | Cupr; HiSG | Cupre; WL |
C. bakeri | Baker/Modoc cypress | Woodland | Cupr; HiSG | Cupre; WL |
C. forbesii | Tecate cypress | Woodland | Cupr; HiSG | Cupre; WL |
C. macrocarpa | Monterey cypress | Woodland | Cupr; HiSG | Cupre; WL |
C. sargentii | Sargent’s cypress | Woodland | Cupr; HiSG | Cupre; WL |
C. macnabiana | MacNab’s cypress | Woodland | Cupr; HiSG | Cupre; WL |
Juniperus spp. | Redcedar/juniper spp. | Cedar/Larch | Cupr; HiSG | Cupre; WL |
J. pinchotii | Pinchot juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. coahuilensis | Redberry juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. flaccida | Drooping juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. ashei | Ashe juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. californica | California juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. deppeana | Alligator juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. occidentalis | Western juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. osteosperma | Utah juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. scopulorum | Rocky Mtn. juniper | Woodland | Cupr; HiSG | Cupre; WL |
J. virginiana var. silcicola | Southern redcedar | Cedar/Larch | Cupr; HiSG | Cupre; WL |
J. virginiana | Easterm redcedar | Cedar/Larch | Cupr; HiSG | Cupre; WL |
J. monosperma | Oneseed juniper | Woodland | Cupr; HiSG | Cupre; WL |
Larix spp. | Larch spp. | Cedar/Larch | Larix | Pinac; WL |
L. laricina | Tamarack | Cedar/Larch | Larix | Pinac; WL |
L. lyallii | Subalpine larch | Cedar/Larch | Larix | Pinac; WL |
L. occidentalis | Western larch | Cedar/Larch | Larix | Pinac; WL |
Calocedrus decurrens | Incense-cedar | Cedar/Larch | Cupr; MedSG | Cupre; WL |
Picea spp. | Spruce spp. | Spruce | Pice; HiSG | Pinac; WL |
P. abies | Norway spruce | Spruce | Pice; HiSG | Pinac; WL |
P. breweriana | Brewer spruce | Spruce | Pice; HiSG | Pinac; WL |
Picea engelmannii | Englemann spruce | Spruce | Pice; LoSG | Pinac; WL |
P. glauca | White spruce | Spruce | Pice; HiSG | Pinac; WL |
P. mariana | Black spruce | Spruce | Pice; HiSG | Pinac; WL |
P. pungens | Blue spruce | Spruce | Pice; HiSG | Pinac; WL |
P. rubens | Red spruce | Spruce | Pice; HiSG | Pinac; WL |
P. sitchensis | Sitka spruce | Spruce | Pice; LoSG | Pinac; WL |
Pinus spp. | Pine spp. | Pine | Pinu; LoSG | Pinac; WL |
P. albicaulis | Whitebark pine | Pine | Pinu; LoSG | Pinac; WL |
P. aristata | Rocky Mtn. bristlecone pine | Pine | Pinu; LoSG | Pinac; WL |
P. attenuata | Knobcone pine | Pine | Pinu; LoSG | Pinac; WL |
P. balfouriana | Foxtail pine | Pine | Pinu; LoSG | Pinac; WL |
P. banksiana | Jack pine | Pine | Pinu; LoSG | Pinac; WL |
P. edulis | Common/two-needle pinyon | Pine | Pinu; HiSG | Pinac; WL |
P. clausa | Sand pine | Pine | Pinu; HiSG | Pinac; WL |
P. contorta | Lodgepole pine | Pine | Pinu; LoSG | Pinac; WL |
P. coulteri | Coulter pine | Pine | Pinu; LoSG | Pinac; WL |
P. echinata | Shortleaf pine | Pine | Pinu; HiSG | Pinac; WL |
P. elliottii | Slash pine | Pine | Pinu; HiSG | Pinac; WL |
P. engelmannii | Apache pine | Pine | Pinu; LoSG | Pinac; WL |
P. flexilis | Limber pine | Pine | Pinu; LoSG | Pinac; WL |
P. strobiformis | Southwestern white pine | Pine | Pinu; LoSG | Pinac; WL |
P. glabra | Spruce pine | Pine | Pinu; LoSG | Pinac; WL |
P. jeffreyi | Jeffrey pine | Pine | Pinu; LoSG | Pinac; WL |
P. lambertiana | Sugar pine | Pine | Pinu; LoSG | Pinac; WL |
P. leiophylla | Chihauhua pine | Pine | Pinu; LoSG | Pinac; WL |
P. monticola | Western white pine | Pine | Pinu; LoSG | Pinac; WL |
P. muricata | Bishop pine | Pine | Pinu; HiSG | Pinac; WL |
P. palustris | Longleaf pine | Pine | Pinu; HiSG | Pinac; WL |
P. ponderosa | Ponderosa pine | Pine | Pinu; LoSG | Pinac; WL |
P. pungens | Table Mountain pine | Pine | Pinu; HiSG | Pinac; WL |
P. radiata | Monterey pine | Pine | Pinu; LoSG | Pinac; WL |
P. resinosa | Red pine | Pine | Pinu; LoSG | Pinac; WL |
P. rigida | Pitch pine | Pine | Pinu; HiSG | Pinac; WL |
P. sabiniana | Gray pine | Pine | Pinu; LoSG | Pinac; WL |
P. serotina | Pond pine | Pine | Pinu; HiSG | Pinac; WL |
P. strobus | Eastern white pine | Pine | Pinu; LoSG | Pinac; WL |
P. sylvestris | Scotch pine | Pine | Pinu; LoSG | Pinac; WL |
P. taeda | Loblolly pine | Pine | Pinu; HiSG | Pinac; WL |
P. virginiana | Viginia pine | Pine | Pinu; HiSG | Pinac; WL |
P. monophylla | Singleleaf pinyon | Pine | Pinu; LoSG | Pinac; WL |
P. discolor | Border pinyon | Pine | Pinu; LoSG | Pinac; WL |
P. arizonica | Arizona pine | Pine | Pinu; LoSG | Pinac; WL |
P. nigra | Austrian pine | Pine | Pinu; LoSG | Pinac; WL |
P. washoensis | Washoe pine | Pine | Pinu; LoSG | Pinac; WL |
P. quadrifolia | Four leaf pine | Pine | Pinu; LoSG | Pinac; WL |
P. torreyana | Torrey pine | Pine | Pinu; LoSG | Pinac; WL |
P. cembroides | Mexican pinyon pine | Pine | Pinu; LoSG | Pinac; WL |
P. remota | Papershell pinyon pine | Pine | Pinu; LoSG | Pinac; WL |
P. longaeva | Great Basin bristlecone pine | Pine | Pinu; LoSG | Pinac; WL |
P. monophylla var. fallax | Arizona pinyon pine | Pine | Pinu; LoSG | Pinac; WL |
P. elliottii var. elliottii | Honduras pine | Pine | Pinu; LoSG | Pinac; WL |
Pseudotsuga spp. | Douglas-fir spp. | Doug Fir | Pseud | Pinac; WL |
P. macrocarpa | Bigcone Douglas-fir | Doug Fir | Pseud | Pinac; WL |
P. menziesii | Douglas-fir | Doug Fir | Pseud | Pinac; WL |
Sequoia sempervirens | Redwood | Cedar/Larch | Cupr; MedSG | Cupre; WL |
Sequoiadendron giganteum | Giant sequoia | Cedar/Larch | Cupr; MedSG | Cupre; WL |
Taxodium spp. | Baldcypress spp. | Cedar/Larch | Cupr; HiSG | Cupre; WL |
T. distichum | Baldcypress | Cedar/Larch | Cupr; HiSG | Cupre; WL |
T. ascendens | Pondcypress | Cedar/Larch | Cupr; HiSG | Cupre; WL |
T. mucronatum | Montezuma baldcypress | Cedar/Larch | Cupr; HiSG | Cupre; WL |
Taxus spp. | Yew spp. | T Fir/Hem | Pseud | |
T. brevifolia | Pacific yew | T Fir/Hem | Pseud | |
T. floridana | Florida yew | T Fir/Hem | Pseud | |
Thuja spp. | Thuja spp. | Cedar/Larch | Cupr; MedSG | Cupre; WL |
T. occidentalis | Northern white-cedar | Cedar/Larch | Cupr; LoSG | Cupre; WL |
T. plicata | Western redcedar | Cedar/Larch | Cupr; MedSG | Cupre; WL |
Torreya spp. | Torreya (nutmeg) spp. | T Fir/Hem | Pseud | |
T. californica | California torreya | T Fir/Hem | Pseud | |
T. taxifolia | Florida torreya | T Fir/Hem | Pseud | |
Tsuga spp. | Hemlock spp. | T Fir/Hem | Tsug; HiSG | Pinac; WL |
T. canadensis | Eastern hemlock | T Fir/Hem | Tsug; LoSG | Pinac; WL |
T. caroliniana | Carolina hemlock | T Fir/Hem | Tsug; HiSG | Pinac; WL |
T. heterophylla | Western hemlock | T Fir/Hem | Tsug; HiSG | Pinac; WL |
T. mertensiana | Mountain hemlock | T Fir/Hem | Tsug; HiSG | Pinac; WL |
Dead conifer | Unknown dead conifer | Pine | Pinu; LoSG | |
Acacia spp. | Acacia spp. | Woodland | Fab/Jug | Fab/Ros; WL |
A. farnesiana | Sweet acacia | Woodland | Fab/Jug | Fab/Ros; WL |
A. greggii | Catclaw acacia | Woodland | Fab/Jug | Fab/Ros; WL |
Acer spp. | Maple spp. | S Maple/Bir | Acer; LoSG | |
A. barbatum | Florida maple | S Maple/Bir | Acer; HiSG | |
A. macrophyllum | Bigleaf maple | S Maple/Bir | Acer; LoSG | |
A. negundo | Boxelder | S Maple/Bir | Acer; LoSG | |
A. nigrum | Black maple | H Maple/Oak | Acer; HiSG | |
A. pensylvanicum | Striped maple | S Maple/Bir | Acer; LoSG | |
A. rubrum | Red maple | S Maple/Bir | Acer; LoSG | |
A. saccharinum | Silver maple | S Maple/Bir | Acer; LoSG | |
A. saccharum | Sugar maple | H Maple/Oak | Acer; HiSG | |
A. spicatum | Mountain maple | S Maple/Bir | Acer; LoSG | |
A. platanoides | Norway maple | S Maple/Bir | Acer; LoSG | |
A. glabrum | Rocky Mtn. maple | Woodland | Acer; LoSG | |
A. grandidentatum | Bigtooth maple | Woodland | Acer; LoSG | |
A. leucoderme | Chalk maple | Mixed HW | Acer; LoSG | |
Aesculus spp. | Buckeye spp. | Mixed HW | Hip/Til | |
A.glabra | Ohio buckeye | Mixed HW | Hip/Til | |
A.flava | Yellow buckeye | Mixed HW | Hip/Til | |
A.californica | California buckeye | Mixed HW | Hip/Til | |
A.glabra var. arguta | Texas buckeye | Mixed HW | Hip/Til | |
A.pavia | Red buckeye | Mixed HW | Hip/Til | |
A.sylvatica | Painted buckeye | Mixed HW | Hip/Til | |
Ailanthus altissima | Ailanthus | Mixed HW | Cor/Eri/Lau/Etc | |
Albizia julibrissin | Mimosa/silktree | Mixed HW | Fab/Jug | Fab/Ros; WL |
Alnus spp. | Alder spp. | Aspen/Alder | Betu; LoSG | |
A. rubra | Red alder | Aspen/Alder | Betu; LoSG | |
A. rhombifolia | White alder | Aspen/Alder | Betu; LoSG | |
A. oblongifolia | Arizona alder | Aspen/Alder | Betu; LoSG | |
A. glutinosa | European alder | Aspen/Alder | Betu; LoSG | |
Amelanchier spp. | Serviceberry spp. | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
A. arborea | Common serviceberry | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
A. sanguinea | Roundleaf serviceberry | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
Arbutus spp. | Madrone spp. | Mixed HW | Cor/Eri/Lau/Etc | |
A. menziesii | Pacific madrone | Mixed HW | Cor/Eri/Lau/Etc | |
A. arizonica | Arizona madrone | Mixed HW | Cor/Eri/Lau/Etc | |
A. xalapensis | Texas madrone | Mixed HW | Cor/Eri/Lau/Etc | |
Asimina triloba | Pawpaw | Mixed HW | Cor/Eri/Lau/Etc | |
Betula spp. | Birch spp. | S Maple/Bir | Betu; Med1SG | |
B. alleghaniensis | Yellow birch | S Maple/Bir | Betu; Med2SG | |
B. lenta | Sweet birch | S Maple/Bir | Betu; HiSG | |
B. nigra | River birch | S Maple/Bir | Betu; Med1SG | |
B. occidentalis | Water birch | S Maple/Bir | Betu; Med2SG | |
B. papyrifera | Paper birch | S Maple/Bir | Betu; Med1SG | |
B. uber | Virginia roundleaf birch | S Maple/Bir | Betu; Med2SG | |
B. utahensis | Northwestern paper birch | S Maple/Bir | Betu; Med2SG | |
B. populifolia | Gray birch | S Maple/Bir | Betu; Med1SG | |
Sideroxylon lanuginosum | Chittamwood/gum bumelia | Mixed HW | Cor/Eri/Lau/Etc | |
Carpinus caroliniana | American hornbeam | Mixed HW | Betu; Med2SG | |
Carya spp. | Hickory spp. | H Maple/Oak | Fab/Jug/Carya | |
C. aquatica | Water hickory | H Maple/Oak | Fab/Jug/Carya | |
C. cordiformis | Bitternut hickory | H Maple/Oak | Fab/Jug/Carya | |
C. glabra | Pignut hickory | H Maple/Oak | Fab/Jug/Carya | |
C. illinoinensis | Pecan | H Maple/Oak | Fab/Jug/Carya | |
C. laciniosa | Shellbark hickory | H Maple/Oak | Fab/Jug/Carya | |
C. myristiciformis | Nutmeg hickory | H Maple/Oak | Fab/Jug/Carya | |
C. ovata | Shagbark hickory | H Maple/Oak | Fab/Jug/Carya | |
C. texana | Black hickory | H Maple/Oak | Fab/Jug/Carya | |
C. alba | Mockernut hickory | H Maple/Oak | Fab/Jug/Carya | |
C. pallida | Sand hickory | H Maple/Oak | Fab/Jug/Carya | |
C. floridana | Scrub hickory | H Maple/Oak | Fab/Jug/Carya | |
C. ovalis | Red hickory | H Maple/Oak | Fab/Jug/Carya | |
C. carolinae-septentrionalis | Southern shagbark hickory | H Maple/Oak | Fab/Jug/Carya | |
Castanea spp. | Chestnut spp. | Mixed HW | Faga; Decid | Fagac; WL |
C. dentata | American chestnut | Mixed HW | Faga; Decid | Fagac; WL |
C. pumila | Allegheny chinkapin | Mixed HW | Faga; Decid | Fagac; WL |
C. pumila var. ozarkensis | Ozark chinkapin | Mixed HW | Faga; Decid | Fagac; WL |
C. mollissima | Chinese chestnut | Mixed HW | Faga; Decid | Fagac; WL |
Chrysolepis chrysophylla | Giant/golden chinkapin | Mixed HW | Faga; Evergrn | Fagac; WL |
Catalpa spp. | Catalpa spp. | Mixed HW | Cor/Eri/Lau/Etc | |
C. bignonioide | Southern catalpa | Mixed HW | Cor/Eri/Lau/Etc | |
C. speciosa | Northern catalpa | Mixed HW | Cor/Eri/Lau/Etc | |
Celtis | Hackberry spp. | Mixed HW | Cor/Eri/Lau/Etc | |
C. laevigata | Sugarberry | Mixed HW | Cor/Eri/Lau/Etc | |
C. occidentalis | Hackberry | Mixed HW | Cor/Eri/Lau/Etc | |
C. laevigata var. reticulata | Netleaf hackberry | Mixed HW | Cor/Eri/Lau/Etc | |
Cercis canadensis | Eastern redbud | Mixed HW | Fab/Jug | Fab/Ros; WL |
Cercocarpus ledifoliu | Curlleaf mountain-mahogany | Woodland | Cor/Eri/Lau/Etc | Fab/Ros; WL |
Cladrastis kentukea | Yellowwood | Mixed HW | Fab/Jug | Fab/Ros; WL |
Cornus spp. | Dogwood spp. | Mixed HW | Cor/Eri/Lau/Etc | |
C. florida | Flowering dogwood | Mixed HW | Cor/Eri/Lau/Etc | |
C. nuttallii | Pacific dogwood | Mixed HW | Cor/Eri/Lau/Etc | |
Crataegus spp. | Hawthorn spp. | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. crusgalli | Cockspur hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. mollis | Downy hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. brainerdii | Brainerd’s hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. calpodendron | Pear hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. chrysocarpa | Fireberry hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. dilatata | Broadleaf hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. flabellata | Fanleaf hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. monogyna | Oneseed hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. pedicellata | Scarlet hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
Eucalyptus spp. | Eucalyptus spp. | Mixed HW | Cor/Eri/Lau/Etc | |
E. globulus | Tasmanian bluegum | Mixed HW | Cor/Eri/Lau/Etc | |
E. camaldulensi | River redgum | Mixed HW | Cor/Eri/Lau/Etc | |
E. grandis | Grand eucalyptus | Mixed HW | Cor/Eri/Lau/Etc | |
E. robusta | Swamp mahogany | Mixed HW | Cor/Eri/Lau/Etc | |
Diospyros spp. | Persimmon spp. | Mixed HW | Cor/Eri/Lau/Etc | |
D. virginiana | Common persimmon | Mixed HW | Cor/Eri/Lau/Etc | |
D. texana | Texas persimmon | Mixed HW | Cor/Eri/Lau/Etc | |
Ehretia anacua | Anacua knockaway | Mixed HW | Cor/Eri/Lau/Etc | |
Fagus grandifolia | American beech | H Maple/Oak | Faga; Decid | Fagac; WL |
Fraxinus spp. | Ash spp. | Mixed HW | Olea; LoSG | |
F. americana | White ash | Mixed HW | Olea; HiSG | |
F. latifolia | Oregon ash | Mixed HW | Olea; LoSG | |
F. nigra | Black ash | Mixed HW | Olea; LoSG | |
F. pennsylvanica | Green ash | Mixed HW | Olea; LoSG | |
F. profunda | Pumpkin ash | Mixed HW | Olea; LoSG | |
F. quadrangulata | Blue ash | Mixed HW | Olea; LoSG | |
F. velutina | Velvet ash | Mixed HW | Olea; LoSG | |
F. caroliniana | Carolina ash | Mixed HW | Olea; LoSG | |
F. texensis | Texas ash | Mixed HW | Olea; LoSG | |
Gleditsia spp. | Honeylocust spp. | Mixed HW | Fab/Jug | Fab/Ros; WL |
G. aquatica | Waterlocust | Mixed HW | Fab/Jug | Fab/Ros; WL |
G. triacanthos | Honeylocust | Mixed HW | Fab/Jug | Fab/Ros; WL |
Gordonia lasianthus | Loblolly-bay | Mixed HW | Cor/Eri/Lau/Etc | |
Ginkgo biloba | Ginkgo | Mixed HW | Cor/Eri/Lau/Etc | |
Gymnocluadus diocicus | Kentucky coffeetree | Mixed HW | Fab/Jug | Fab/Ros; WL |
Halesia spp. | Silverbell spp. | Mixed HW | Cor/Eri/Lau/Etc | |
H. carolina | Carolina silverbell | Mixed HW | Cor/Eri/Lau/Etc | |
H. diptera | Two-wing silverbell | Mixed HW | Cor/Eri/Lau/Etc | |
H. parviflora | Little silverbell | Mixed HW | Cor/Eri/Lau/Etc | |
Ilex opaca | American holly | Mixed HW | Cor/Eri/Lau/Etc | |
Juglans spp. | Walnut spp. | Mixed HW | Fab/Jug | |
J. cinerea | Butternut | Mixed HW | Fab/Jug | |
J. nigra | Black walnut | Mixed HW | Fab/Jug | |
J. hindsii | No. California black walnut | Mixed HW | Fab/Jug | |
J. californica | So. California black walnut | Mixed HW | Fab/Jug | |
J. microcarpa | Texas walnut | Mixed HW | Fab/Jug | |
J. major | Arizona walnut | Mixed HW | Fab/Jug | |
Liquidambar styraciflua | Sweetgum | Mixed HW | Hama | |
Liriodendron tulipifera | Yellow poplar | Mixed HW | Magno | |
Lithocarpus densiflorus | Tanoak | Mixed HW | Faga; Evergrn | Fagac; WL |
Maclura pomifera | Osage orange | Mixed HW | Cor/Eri/Lau/Etc | |
Magnolia spp. | Magnolia spp. | Mixed HW | Magno | |
M. acuminata | Cucumbertree | Mixed HW | Magno | |
M. grandiflora | Southern magnolia | Mixed HW | Magno | |
M. virginiana | Sweeetbay | Mixed HW | Magno | |
M. macrophylla | Bigleaf magnolia | Mixed HW | Magno | |
M. fraseri | Mountain/Frasier magnolia | Mixed HW | Magno | |
M. pyramidata | Pyramid magnolia | Mixed HW | Magno | |
M. tripetala | Umbrella magnolia | Mixed HW | Magno | |
Malus spp. | Apple spp. | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
M. fusca | Oregon crab apple | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
M. angustifolia | Southern crabapple | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
M. coronaria | Sweet crabapple | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
M. ioensi | Prairie crabapple | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
Morus spp. | Mulberry spp. | Mixed HW | Cor/Eri/Lau/Etc | |
M. alba | White mulberry | Mixed HW | Cor/Eri/Lau/Etc | |
M. rubra | Red mulberry | Mixed HW | Cor/Eri/Lau/Etc | |
M. microphyll | Texas mulberry | Mixed HW | Cor/Eri/Lau/Etc | |
M. nigra | Black mulberry | Mixed HW | Cor/Eri/Lau/Etc | |
Nyssa spp. | Tupelo spp. | Mixed HW | Cor/Eri/Lau/Etc | |
N. aquatica | Water tupelo | Mixed HW | Cor/Eri/Lau/Etc | |
N. ogeche | Ogeechee tupelo | Mixed HW | Cor/Eri/Lau/Etc | |
N. sylvatica | Blackgum | Mixed HW | Cor/Eri/Lau/Etc | |
N. biflora | Swamp tupelo | Mixed HW | Cor/Eri/Lau/Etc | |
Ostrya virginiana | Eastern hophornbeam | Mixed HW | Betu; HiSG | |
Oxydendrum arboreum | Sourwood | Mixed HW | Cor/Eri/Lau/Etc | |
Paulownia tomentosa | Paulownia/empress tree | Mixed HW | Cor/Eri/Lau/Etc | |
Persea spp. | Bay spp. | Mixed HW | Cor/Eri/Lau/Etc | |
Persea borbonia | Redbay | Mixed HW | Cor/Eri/Lau/Etc | |
Planera aquatica | Water elm/planetree | Mixed HW | Cor/Eri/Lau/Etc | |
Platanus spp. | Sycamore spp. | Mixed HW | Cor/Eri/Lau/Etc | |
P. racemosa | California sycamore | Mixed HW | Cor/Eri/Lau/Etc | |
P. occidentalis | American sycamore | Mixed HW | Cor/Eri/Lau/Etc | |
P. wrightii | Arizona sycamore | Mixed HW | Cor/Eri/Lau/Etc | |
Populus spp. | Cottonwood/poplar spp. | Aspen/Alder | Sali; HiSG | |
P. balsamifera | Balsam poplar | Aspen/Alder | Sali; LoSG | |
P. deltoides | Eastern cottonwood | Aspen/Alder | Sali; HiSG | |
P. grandidentata | Bigtooth aspen | Aspen/Alder | Sali; HiSG | |
P. heterophylla | Swamp cottonwood | Aspen/Alder | Sali; HiSG | |
P. deltoides | Plains cottonwood | Aspen/Alder | Sali; HiSG | |
P. tremuloides | Quaking aspen | Aspen/Alder | Sali; HiSG | |
P. balsamifera | Black cottonwood | Aspen/Alder | Sali; LoSG | |
P. fremontii | Fremont cottonwood | Aspen/Alder | Sali; HiSG | |
P. angustifolia | Narrlowleaf cottonwood | Aspen/Alder | Sali; HiSG | |
P. alba | Silver poplar | Aspen/Alder | Sali; HiSG | |
P. nigra | Lombardy poplar | Aspen/Alder | Sali; HiSG | |
Prosopis spp. | Mesquite spp. | Woodland | Fab/Jug | Fab/Ros; WL |
P. glandulosa | Honey mesquite | Woodland | Fab/Jug | Fab/Ros; WL |
P. velutina | Velvet mesquite | Woodland | Fab/Jug | Fab/Ros; WL |
P. pubescens | Screwbean mesquite | Woodland | Fab/Jug | Fab/Ros; WL |
Prunus spp. | Cherry/plum spp. | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. pensylvanica | Pin cherry | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. serotina | Black cherry | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. virginiana | Chokecherry | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. persica | Peach | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. nigra | Canada plum | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. americana | American plum | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. emarginata | Bitter cherry | Woodland | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. alleghaniensis | Allegheny plum | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. angustifolia | Chickasaw plum | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. avium | Sweet cherry (domestic) | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. cerasus | Sour cherry (domestic) | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. domestica | European plum (domestic) | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
P. mahaleb | Mahaleb cherry (domestic) | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
Quercus spp. | Oak spp. | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. agrifolia | California live oak | H Maple/Oak | Faga; Evergrn | Fagac; WL |
Q. alba | White oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. arizonica | Arizona white oak | Woodland | Faga; Decid | Fagac; WL |
Q. bicolor | Swamp white oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. chrysolepis | Canyon live oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. coccinea | Scarlet oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. douglasii | Blue oak | H Maple/Oak | Faga; Evergrn | Fagac; WL |
Q. sinuata var. sinuata | Durand oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. ellipsoidalis | Northern pin oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. emoryi | Emory oak | Woodland | Faga; Decid | Fagac; WL |
Q. engelmannii | Englemann oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. falcata | Southern red oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. pagoda | Cherrybark oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. gambelii | Gambel oak | Woodland | Faga; Decid | Fagac; WL |
Q. garryana | Oregon white oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. ilicifolia | Scrub oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. imbricaria | Shingle oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. kelloggii | California black oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. laevis | Turkey oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. laurifolia | Laurel oak | H Maple/Oak | Faga; Evergrn | Fagac; WL |
Q. lobata | California white oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. lyrata | Overcup oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. macrocarpa | Bur oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. marilandica | Blackjack oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. michauxi | Swamp chestnut oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. muehlenbergii | Chinkapin oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. nigra | Water oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. texana | Texas red oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. oblongifolia | Mexican blue oak | Woodland | Faga; Decid | Fagac; WL |
Q. palustris | Pin oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. phellos | Willow oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. prinus | Chestnut oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. rubra | Northern red oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. shumardii | Shumard oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. stellata | Post oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. simili | Delta post oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. velutina | Black oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. virginiana | Live oak | H Maple/Oak | Faga; Evergrn | Fagac; WL |
Q. wislizeni | Interier live oak | H Maple/Oak | Faga; Evergrn | Fagac; WL |
Q. margarettiae | Dwarf post oak | H Maple/Oak | Faga; Evergrn | Fagac; WL |
Q. minima | Dwarf live oak | H Maple/Oak | Faga; Evergrn | Fagac; WL |
Q. incana | Bluejack oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. hypoleucoides | Silverleaf oak | Woodland | Faga; Decid | Fagac; WL |
Q. oglethorpensis | Oglethorpe oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. prinoides | Dwarf chinkapin oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. grisea | Gray oak | Woodland | Faga; Decid | Fagac; WL |
Q. rugosa | Netleaf oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. gracilliformis | Chisos oak | Woodland | Faga; Decid | Fagac; WL |
Amyris elemifera | Sea torchwood | Mixed HW | Cor/Eri/Lau/Etc | |
Annona glabra | Pond apple | Mixed HW | Cor/Eri/Lau/Etc | |
Bursera simaruba | Gumbo limbo | Mixed HW | Cor/Eri/Lau/Etc | |
Casuarina spp. | Sheoak spp. | Mixed HW | Cor/Eri/Lau/Etc | |
C. glauca | Gray sheoak | Mixed HW | Cor/Eri/Lau/Etc | |
C. lepidophloia | Belah | Mixed HW | Cor/Eri/Lau/Etc | |
Cinnamomum camphora | Camphortree | Mixed HW | Cor/Eri/Lau/Etc | |
Citharexylum fruticosum | Florida fiddlewood | Mixed HW | Cor/Eri/Lau/Etc | |
Citrus spp. | Citrus spp. | Mixed HW | Cor/Eri/Lau/Etc | |
Coccoloba diversifolia | Tietongue/pigeon plum | Mixed HW | Cor/Eri/Lau/Etc | |
Colubrina elliptica | Soldierwood | Mixed HW | Cor/Eri/Lau/Etc | |
Cordia sebestena | Longleaf geigertree | Mixed HW | Cor/Eri/Lau/Etc | |
Cupaniopsis anacardioides | Carrotwood | Mixed HW | Cor/Eri/Lau/Etc | |
Condalia hookeri | Bluewood | Woodland | Cor/Eri/Lau/Etc | |
Ebenopsis ebano | Blackbead ebony | Woodland | Fab/Jug | Fab/Ros; WL |
Leucaena pulverulenta | Great leadtree | Woodland | Fab/Jug | Fab/Ros; WL |
Sophora affinis | Texas sophora | Woodland | Fab/Jug | Fab/Ros; WL |
Eugenia rhombea | Red stopper | Mixed HW | Cor/Eri/Lau/Etc | |
Exothea paniculata | Butterbough/inkwood | Mixed HW | Cor/Eri/Lau/Etc | |
Ficus aurea | Florida strangler fig | Mixed HW | Cor/Eri/Lau/Etc | |
Ficus citrifolia | Banyantree/shortleaf fig | Mixed HW | Cor/Eri/Lau/Etc | |
Guapira discolo | Beeftree/longleaf blolly | Mixed HW | Cor/Eri/Lau/Etc | |
Hippomane mancinella | Manchineel | Mixed HW | Cor/Eri/Lau/Etc | |
Lysiloma latisiliquum | False tamarind | Mixed HW | Fab/Jug | Fab/Ros; WL |
Mangifera indica | Mango | Mixed HW | Cor/Eri/Lau/Etc | |
Metopium toxiferum | Florida poisontree | Mixed HW | Cor/Eri/Lau/Etc | |
Piscidia piscipula | Fishpoison tree | Mixed HW | Fab/Jug | Fab/Ros; WL |
Schefflera actinophylla | Octopus tree/schefflera | Mixed HW | Cor/Eri/Lau/Etc | |
Sideroxylon foetidissimum | False mastic | Mixed HW | Cor/Eri/Lau/Etc | |
Sideroxylon salicifolium | White bully/willow bustic | Mixed HW | Cor/Eri/Lau/Etc | |
Simarouba glauca | Paradisetree | Mixed HW | Cor/Eri/Lau/Etc | |
Syzygium cumini | Java plum | Mixed HW | Cor/Eri/Lau/Etc | |
Tamarindus indica | Tamarind | Mixed HW | Fab/Jug | Fab/Ros; WL |
Robinia pseudoacacia | Black locust | Mixed HW | Fab/Jug | Fab/Ros; WL |
Robinia neomexicana | New Mexico locust | Woodland | Fab/Jug | Fab/Ros; WL |
Acoelorraphe wrightii | Everglades palm | Mixed HW | Cor/Eri/Lau/Etc | |
Coccothrinax argentata | Florida silver palm | Mixed HW | Cor/Eri/Lau/Etc | |
Cocos nucifera | Coconut palm | Mixed HW | Cor/Eri/Lau/Etc | |
Roystonea spp. | Royal palm spp. | Mixed HW | Cor/Eri/Lau/Etc | |
Sabal Mexicana | Mexican palmetto | Mixed HW | Cor/Eri/Lau/Etc | |
Sabal palmetto | Cabbage palmetto | Mixed HW | Cor/Eri/Lau/Etc | |
Thrinax morrisii | Key thatch palm | Mixed HW | Cor/Eri/Lau/Etc | |
Thrinax radiata | Florida thatch palm | Mixed HW | Cor/Eri/Lau/Etc | |
Arecaceae | Other palms | Mixed HW | Cor/Eri/Lau/Etc | |
Sapindus saponaria | Western soapberry | Mixed HW | Cor/Eri/Lau/Etc | |
Salix spp. | Willow spp. | Aspen/Alder | Sali; HiSG | |
S. amygdaloides | Peachleaf willow | Aspen/Alder | Sali; HiSG | |
S. nigra | Black willow | Aspen/Alder | Sali; HiSG | |
S. bebbiana | Bebb willow | Aspen/Alder | Sali; HiSG | |
S. bonplandiana | Bonpland willow | Aspen/Alder | Sali; HiSG | |
S. caroliniana | Coastal plain willow | Aspen/Alder | Sali; HiSG | |
S. pyrifolia | Balsam willow | Aspen/Alder | Sali; HiSG | |
S. alba | White willow | Aspen/Alder | Sali; HiSG | |
S. scouleriana | Scouder’s willow | Aspen/Alder | Sali; HiSG | |
S. sepulcralis | Weeping willow | Aspen/Alder | Sali; HiSG | |
Sassafras albidum | Sassafrass | Mixed HW | Cor/Eri/Lau/Etc | |
Sorbus spp. | Mountain ash spp. | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
S. americana | American mountain ash | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
S. aucuparia | European mountain ash | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
S. decora | Northern mountain ash | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
Swietenia mahagoni | West Indian mahogany | Mixed HW | Cor/Eri/Lau/Etc | |
Tilia spp. | Basswood spp. | Mixed HW | Hip/Til | |
T. americana | American basswood | Mixed HW | Hip/Til | |
T. americana var. heterophylla | White basswood | Mixed HW | Hip/Til | |
T. americana var. caroliniana | Carolina basswood | Mixed HW | Hip/Til | |
Ulmus spp. | Elm spp. | Mixed HW | Cor/Eri/Lau/Etc | |
U. alata | Winged elm | Mixed HW | Cor/Eri/Lau/Etc | |
U. americana | American elm | Mixed HW | Cor/Eri/Lau/Etc | |
U. crassifolia | Cedar elm | Mixed HW | Cor/Eri/Lau/Etc | |
U. pumila | Siberian elm | Mixed HW | Cor/Eri/Lau/Etc | |
U. rubra | Slippery elm | Mixed HW | Cor/Eri/Lau/Etc | |
U. serotina | September elm | Mixed HW | Cor/Eri/Lau/Etc | |
U. thomasii | Rock elm | Mixed HW | Cor/Eri/Lau/Etc | |
Umbellularia californica | California laurel | Mixed HW | Cor/Eri/Lau/Etc | |
Yucca brevifolia | Joshua tree | Mixed HW | Cor/Eri/Lau/Etc | |
Avicennia germinan | Black mangrove | Mixed HW | Cor/Eri/Lau/Etc | |
Conocarpus erectus | Button mangrove | Mixed HW | Cor/Eri/Lau/Etc | |
Laguncularia racemosa | White mangrove | Mixed HW | Cor/Eri/Lau/Etc | |
Rhizophora mangle | American mangrove | Mixed HW | Cor/Eri/Lau/Etc | |
Olneya tesota | Desert ironwood | Woodland | Fab/Jug | Fab/Ros; WL |
Tamarix spp. | Saltcedar | Mixed HW | Cor/Eri/Lau/Etc | |
Melaleuca quinquenervia | Melaleuca | Mixed HW | Cor/Eri/Lau/Etc | |
Melia azedarach | Chinaberry | Mixed HW | Cor/Eri/Lau/Etc | |
Triadica sebifera | Chinese tallowtree | Mixed HW | Cor/Eri/Lau/Etc | |
Vernicia fordii | Tungoil tree | Mixed HW | Cor/Eri/Lau/Etc | |
Cotinus obovatus | Smoketree | Mixed HW | Cor/Eri/Lau/Etc | |
Elaeagnus angustifolia | Russian olive | Mixed HW | Cor/Eri/Lau/Etc | |
Tree broadleaf | Unknown dead hardwood | Mixed HW | Cor/Eri/Lau/Etc | |
Tree unknown | Unknown live tree | Mixed HW | Cor/Eri/Lau/Etc | |
C. phaenopyrum | Washington hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. succulenta | Fleshy hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
C. uniflora | Dwarf hawthorn | Mixed HW | Cor/Eri/Lau/Etc | Fab/Ros; WL |
F. berlandieriana | Berlandier ash | Mixed HW | Olea; LoSG | |
Persea americana | Avocado | Mixed HW | Cor/Eri/Lau/Etc | |
Ligustrum sinense | Chinese privet | Mixed HW | Olea; HiSG | |
Q. gravesii | Graves oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. polymorpha | Mexican white oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. buckleyi | Buckley oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Q. laceyi | Lacey oak | H Maple/Oak | Faga; Decid | Fagac; WL |
Cordia boissieri | Anacahuita Texas olive | Mixed HW | Cor/Eri/Lau/Etc | |
Tamarix aphylla | Athel tamarisk | Mixed HW | Cor/Eri/Lau/Etc |
- 1.
Overall, follow the placement of taxa as suggested within the manuscript (i.e., as in Tables 2, 3, 4, and Figs. 2, 3, and 4).
- 2.
If a tree record is one of the five families (of Table 4) and the tree diameter is measured as diameter at root collar then one of the Table 4 woodland equations applies. Otherwise, if one of the five (Table 4) families and diameter is dbh then use the appropriate equation from Tables 2 or 3. If not one of the five Table 4 families but tree diameter is provided as a root collar measurement, then convert drc to dbh following information provided in Fig. 1 before applying a Table 2 or 3 equation.
- 3.
The calculations for the woodland (Table 4) Cupressaceae (“Cupre; WL”) uses the “2nd juniper” equation from footnote #2 in Table 5.
- 4.
The Fabaceae/Juglandaceae split into the two groups—“Fab/Jug/Carya” and “Fab/Jug”—is according to the genus Carya versus all others (i.e., not-Carya ).
- 5.
Fagaceae’s deciduous/evergreen split—“Faga; Decid” and “Faga; Evergrn”—sets deciduous as the default. The Fagaceae allocated to evergreen are those five species explicitly listed as evergreen in Table 3 and those identified as evergreen from the USDA PLANTS database [19], which currently includes the addition of three live oak species.
- 6.
The 6-family general equation at the middle of page 136 (in Table 3 of Chojnacky et al. [12])—“Cor/Eri/Lau/Etc”—is assigned trees by family from 3 sources: (a) the six families listed in Table 3; (b) the five additional families noted in the Fig. 3 caption, and (c) any additional formerly unassigned hardwood species.
- 7.
Roots—the Chojnacky estimates use both of the belowground root equations of Table 6 (the sum of the two is generally equivalent to the original Jenkins root component). Note these are dbh-based, so a drc tree should first convert drc-to-dbh according to Fig. 1. Also note, all other (other than root) components of the original Jenkins et al. [1] are applicable here.
Identifying equivalence between the alternate biomass estimates
Tests of equivalence of the plot level (tonnes carbon per hectare) representation of the Jenkins and Chojnacky groups are included principally as guidance as to where the choice of biomass equations may matter. The analysis does not address relative accuracy of the two alternatives. Specifically, we focused on equivalence tests of the mean difference between the two estimates at the plot, or stand, level according to region and forest type groups. While these are species (group) level equations, any practical effect (of interest) is at plot to landscape to national (carbon reporting) levels. Equivalence tests are appropriate where the questions are more directly “are the groups similar, or effectively the same?” and not so much “are they different?” [20, 21]. This distinction follows from the idea that failure to reject a null hypothesis of no difference between populations does not necessarily indicate that the null hypothesis is true. The essential characteristic of an equivalence test is that the null hypothesis is stated such that the two populations are different [22, 23] which can be viewed as the reverse of the more common approach to hypothesis testing. The specific measure, or threshold, of where two populations can be considered equivalent versus different is set by researchers and a conclusion of not-different, or equivalent, results from rejecting the null hypothesis (that the two are different).
Equivalence tests presented here are paired-sample tests [24, 25] because each sample is based on estimates from each of the Chojnacky and Jenkins groups. Our test statistic is the difference between estimates (Chojnacky minus Jenkins), and we set “equivalence” as a mean difference less than 5 % of the Jenkins-based estimate. Putting our test in terms of the null and alternative hypotheses following the format of publications describing this approach [22, 24], we have:
Null, H_{0}: (Chojnacky-Jenkins) <−5 % Jenkins or (Chojnacky-Jenkins) >5 Jenkins
and
Alternative, H_{1}: −5 % Jenkins ≤ (Chojnacky-Jenkins) ≤5 % Jenkins
We use the two one-sided tests (TOST) of our two-part null hypothesis that the plot-level difference was greater than 5 % of the Jenkins value and set α = 0.05—one test that the mean difference is less than minus 5 % of the Jenkins estimate, and one test that the mean difference is greater than 5 % of the Jenkins estimate. Within an application of the TOST where α is set to 0.05, a one-step approach to accomplish the TOST result is establish a 2-sided 90 % confidence interval for the test statistic; if this falls entirely within the prescribed interval then the two populations can be considered equivalent [26]. We also extended the level of “equivalence” to within 10 % of the Jenkins-based estimates for some analyses in order to look for more general trends, or broad agreement between the two approaches.
Our equivalence tests are based on the paired estimates of carbon tonnes per hectare on the single-condition forested plots variously classified according to regions described in Fig. 1, forest type-groups listed in Table 1, or stand size class as in Fig. 3 (see [13] for additional details about these classifications). The distribution of the test statistic (mean difference) was obtained from resampling with replacement [27] ten thousand times, with a mean value determined for each sample. The number of plots available varied depending on the classification (Table 1; Fig. 3). We did not test for equivalence if fewer than 30 plots were available, and if over 2000 plots were available we randomly selected 2000 for resampling. The choice of 2000 is based on preliminary analysis of these data that showed the confidence interval from resampling converge with percentiles obtained directly from the distribution of the large number of sample plots, usually well below 1000; the 2000 is simply a round number well beyond this convergence without getting too computationally intense. The 90 % confidence interval (the same as the 95 % interval of TOST) obtained for the distribution of the mean difference is according to a bias corrected and accelerated percentile method [28, 29]. Note that our tests for equivalence are based on comparing this confidence interval to the ±5 % of the corresponding Jenkins based estimate. Table 1 provides the estimates from the two approaches, with the equivalence test results indicated with asterisks. Similarly, the equivalence test results in Fig. 3 are not in the tonnes per hectare of the resampled values and the confidence interval, they are represented as percentage of Jenkins estimates—for this, equivalence is established if the entire confidence interval is within the zero side of the respective 5 %.
Declarations
Authors’ contributions
Design and analysis was split equally between JS and CH; JS was responsible for coding and calculations, CH developed the figures and tables, and writing was equally divided between JS and CH. All authors read and approved the final manuscript.
Acknowledgements
The authors gratefully acknowledge the helpful feedback from Linda Heath and William Leak on the draft manuscript. We would also like to thank the anonymous reviewers for their time and comments.
Competing interests
The authors declare that they have no competing interests.
Open AccessThis article is distributed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted use, distribution, and reproduction in any medium, provided you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made.
Authors’ Affiliations
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